Ownership Submission
FORM 3
UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

INITIAL STATEMENT OF BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940
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(Print or Type Responses)
1. Name and Address of Reporting Person *
  Polaris Venture Management Co. V, L.L.C.
2. Date of Event Requiring Statement (Month/Day/Year)
09/30/2013
3. Issuer Name and Ticker or Trading Symbol
FATE THERAPEUTICS INC [FATE]
(Last)
(First)
(Middle)
C/O POLARIS VENTURE PARTNERS, 1000 WINTER STREET, SUITE 3350
4. Relationship of Reporting Person(s) to Issuer
(Check all applicable)
_____ Director __X__ 10% Owner
_____ Officer (give title below) _____ Other (specify below)
5. If Amendment, Date Original Filed(Month/Day/Year)
(Street)

WALTHAM, MA 02451
6. Individual or Joint/Group Filing(Check Applicable Line)
___ Form filed by One Reporting Person
_X_ Form filed by More than One Reporting Person
(City)
(State)
(Zip)
Table I - Non-Derivative Securities Beneficially Owned
1.Title of Security
(Instr. 4)
2. Amount of Securities Beneficially Owned
(Instr. 4)
3. Ownership Form: Direct (D) or Indirect (I)
(Instr. 5)
4. Nature of Indirect Beneficial Ownership
(Instr. 5)

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly. SEC 1473 (7-02)
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.

Table II - Derivative Securities Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivative Security
(Instr. 4)
2. Date Exercisable and Expiration Date
(Month/Day/Year)
3. Title and Amount of Securities Underlying Derivative Security
(Instr. 4)
4. Conversion or Exercise Price of Derivative Security 5. Ownership Form of Derivative Security: Direct (D) or Indirect (I)
(Instr. 5)
6. Nature of Indirect Beneficial Ownership
(Instr. 5)
Date Exercisable Expiration Date Title Amount or Number of Shares
Series A Convertible Preferred Stock   (1)   (1) Common Stock 651,806 $ (1) I By Polaris Venture Partners V, L.P. (3)
Series A Convertible Preferred Stock   (1)   (1) Common Stock 12,704 $ (1) I By Polaris Venture Partners Entrepreneurs' Fund V, L.P. (4)
Series A Convertible Preferred Stock   (1)   (1) Common Stock 4,465 $ (1) I By Polaris Venture Partners Founders' Fund V, L.P. (5)
Series A Convertible Preferred Stock   (1)   (1) Common Stock 6,518 $ (1) I By Polaris Venture Partners Special Founders' Fund V, L.P. (6)
Series B Convertible Preferred Stock   (2)   (2) Common Stock 255,950 $ (2) I By Polaris Venture Partners V, L.P. (3)
Series B Convertible Preferred Stock   (2)   (2) Common Stock 4,988 $ (2) I By Polaris Venture Partners Entrepreneurs' Fund V, L.P. (4)
Series B Convertible Preferred Stock   (2)   (2) Common Stock 1,753 $ (2) I By Polaris Venture Partners Founders' Fund V, L.P. (5)
Series B Convertible Preferred Stock   (2)   (2) Common Stock 2,560 $ (2) I By Polaris Venture Partners Special Founders' Fund V, L.P. (6)
Series C Convertible Preferred Stock   (1)   (1) Common Stock 550,653 $ (1) I By Polaris Venture Partners V, L.P. (3)
Series C Convertible Preferred Stock   (1)   (1) Common Stock 10,732 $ (1) I By Polaris Venture Partners Entrepreneurs' Fund V, L.P. (4)
Series C Convertible Preferred Stock   (1)   (1) Common Stock 3,772 $ (1) I By Polaris Venture Partners Founders' Fund V, L.P. (5)
Series C Convertible Preferred Stock   (1)   (1) Common Stock 5,506 $ (1) I By Polaris Venture Partners Special Founders' Fund V, L.P. (6)

Reporting Owners

Reporting Owner Name / Address Relationships
Director 10% Owner Officer Other
Polaris Venture Management Co. V, L.L.C.
C/O POLARIS VENTURE PARTNERS
1000 WINTER STREET, SUITE 3350
WALTHAM, MA 02451
    X    
Polaris Venture Partners V, L.P.
1000 WINTER STREET, SUITE 3350
WALTHAM, MA 02451
    X    
Polaris Venture Partners Entrepreneurs' Fund V, L.P.
1000 WINTER STREET, SUITE 3350
WALTHAM, MA 02451
    X    
Polaris Venture Partners Founders' Fund V, L.P.
1000 WINTER STREET, SUITE 3350
WALTHAM, MA 02451
    X    
Polaris Venture Partners Special Founders' Fund V, L.P.
1000 WINTER STREET, SUITE 3350
WALTHAM, MA 02451
    X    

Signatures

/s/ Jonathan Flint, Authorized Signatory for Polaris Venture Management Co. V, L.L.C. 09/30/2013
**Signature of Reporting Person Date

/s/ Jonathan Flint, Authorized Signatory for Polaris Venture Partners V, L.P. 09/30/2013
**Signature of Reporting Person Date

/s/ Jonathan Flint, Authorized Signatory for Polaris Venture Partners Entrepreneurs' Fund V, L.P. 09/30/2013
**Signature of Reporting Person Date

/s/ Jonathan Flint, Authorized Signatory for Polaris Venture Partners Founders' Fund V, L.P. 09/30/2013
**Signature of Reporting Person Date

/s/ Jonathan Flint, Authorized Signatory for Polaris Venture Partners Special Founders' Fund V, L.P. 09/30/2013
**Signature of Reporting Person Date

Explanation of Responses:

* If the form is filed by more than one reporting person, see Instruction 5(b)(v).
** Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1) The aggregate number of shares of Issuer's Preferred Stock held by the Reporting Person converts into Common Stock immediately prior to the closing of the Issuer's initial public offering and has no expiration date. These shares convert into Common Stock on a 1-for-1 basis.
(2) The aggregate number of shares of Issuer's Preferred Stock held by the Reporting Person converts into Common Stock immediately prior to the closing of the Issuer's initial public offering and has no expiration date. These shares convert into Common Stock on approximately a 1-for-1.15 basis.
(3) These shares are owned directly by Polaris Venture Partners V, L.P., whose sole general partner is Polaris Venture Management Co. V, LLC ("Polaris Management"). The members of North Star Venture Management 2010 LLC are also members of Polaris Management, and as members of the general partner, they may be deemed to share voting and investment power of such shares. The Reporting Person is a member of Polaris Management and has shared voting and investment power over such shares and may be deemed the indirect beneficial owner of such shares. The Reporting Person disclaims beneficial ownership of these securities, except to the extent of his pecuniary interest therein, and this report shall not be deemed an admission that the Reporting Person is the beneficial owner of such securities for Section 16 or any other purpose.
(4) These shares are owned directly by Polaris Venture Partners Entrepreneurs' Fund V, L.P., whose sole general partner is Polaris Management. The members of North Star Venture Management 2010 LLC are also members of Polaris Management, and as members of the general partner, they may be deemed to share voting and investment power of such shares. The Reporting Person is a member of Polaris Management and has shared voting and investment power over such shares and may be deemed the indirect beneficial owner of such shares. The Reporting Person disclaims beneficial ownership of these securities, except to the extent of his pecuniary interest therein, and this report shall not be deemed an admission that the Reporting Person is the beneficial owner of such securities for Section 16 or any other purpose.
(5) These shares are owned directly by Polaris Venture Partners Founders' Fund V, L.P., whose sole general partner is Polaris Management. The members of North Star Venture Management 2010 LLC are also members of Polaris Management, and as members of the general partner, they may be deemed to share voting and investment power of such shares. The Reporting Person is a member of Polaris Management and has shared voting and investment power over such shares and may be deemed the indirect beneficial owner of such shares. The Reporting Person disclaims beneficial ownership of these securities, except to the extent of his pecuniary interest therein, and this report shall not be deemed an admission that the Reporting Person is the beneficial owner of such securities for Section 16 or any other purpose.
(6) These shares are owned directly by Polaris Venture Partners Special Founders' Fund V, L.P., whose sole general partner is Polaris Management. The members of North Star Venture Management 2010 LLC are also members of Polaris Management, and as members of the general partner, they may be deemed to share voting and investment power of such shares. The Reporting Person is a member of Polaris Management and has shared voting and investment power over such shares and may be deemed the indirect beneficial owner of such shares. The Reporting Person disclaims beneficial ownership of these securities, except to the extent of his pecuniary interest therein, and this report shall not be deemed an admission that the Reporting Person is the beneficial owner of such securities for Section 16 or any other purpose.

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